Life cycle of Plasmodium falciparum
The life cycle of all Plasmodium species is complex. Infection in
humans begins with the bite of an infected female Anopheline
mosquito. Sporozoites released from the salivary glands of the
mosquito enter the bloodstream during feeding quickly invade liver cells
(hepatocytes). Sporozoites are cleared from the circulation within 30 minutes.
During the next 14 days in the case of P. falciparum, the liver-stage
parasites differentiate and undergo asexual multiplication resulting in
tens of thousands of merozoites which burst from the hepatoctye.
Individual merozoites invade red blood cells (erythrocytes) and undergo
an additional round of multiplication producing 12- 16 merozoites within
a schizont. The length of this erythrocytic stage of the parasite
life cycle depends on the parasite species: 48 hours for P. falciparum,
P. vivax, and P. ovale and 72 hours for P. malariae. The clinical manifestations
of malaria, fever and chills, are associated with the synchronous rupture
of the infected erythrocyte. The released merozoites go on to invade additional
erythrocytes. Not all of the merozoites divide into schizonts, some differentiate
into sexual forms, male and female gametocytes. These gametocytes
are taken up by a female anophylean mosquito during a blood meal. Within
the mosquito midgut, the male gametocyte undergoes a rapid nuclear division,
producing 8 flagellated microgamtes which fertilize the female macrogamete.
The resulting ookinete traverses the mosquito gut wall and encysts
on the exterior of the gut wall as a oocyst. Soon the oocyst ruptures,
releasing hundreds of sporozoites into the mosquito body cavity where they
eventually migrate to the mosquito salivary gland.
The Mosquito vector, female Anopheline mosquito